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Naive and idealistic Jefferson Smith, leader of the Boy Rangers, is appointed to the United States Senate by the puppet governor of his state. He soon discovers, upon going to Washington, many shortcomings of the political process as his earnest goal of a national boys' camp leads to a conflict with the state political boss.
As a parting shot, fired reporter Ann Mitchell prints a fake letter from unemployed "John Doe," who threatens suicide in protest of social ills. The paper is forced to rehire Ann and hires John Willoughby to impersonate "Doe." Ann and her bosses cynically milk the story for all it's worth, until the made-up "John Doe" philosophy starts a whole political movement.
Gunslinger Annie Oakley romances fellow sharpshooter Frank Butler as they travel with Buffalo Bill's Wild West Show.
Alice, the only relatively normal member of the eccentric Sycamore family, falls in love with Tony Kirby, but his wealthy banker father and snobbish mother strongly disapprove of the match. When the Kirbys are invited to dinner to become better acquainted with their future in-laws, things don't turn out the way Alice had hoped.