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Sonic's arch nemesis Dr. Robotnik has been banished from the Land of Darkness by an evil Metal Robotnik. The devious doctor tells Sonic that the Robot Generator has been sabotaged and will blow Planet Freedom to kingdom come. But it's not until the President's beautiful daughter Sara turns on the charm that Sonic springs into action.
Di Gi Charat (a.k.a. Dejiko) - along with Petit Charat (a.k.a. Puchiko) and Gema - travels to Earth as part of her training to become a full-fledged princess. They crash on a small town in Japan, where they meet the Omocha brothers (who spend most of their time thinking how cute Puchiko is) and Mr. & Mrs. Ankoro (an elderly couple that makes Japanese sweets).
Erika, the princess of the Magic Kingdom has come to Earth in order to find a human girl who looks just like her. That girl turns out to be Himeko Nonohara, a tomboy's tomboy. Erika must give Himeko a magical item she has created in order to prove her worth as a successor to the crown. Himeko must test this item, a hair ribbon that allows her to transform into any other person she sees, to see if it is worthy. The series follows Himeko's adventures and her budding romance with Daichi, the boy who discovers her secret.