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Lightning McQueen, a hotshot rookie race car driven to succeed, discovers that life is about the journey, not the finish line, when he finds himself unexpectedly detoured in the sleepy Route 66 town of Radiator Springs. On route across the country to the big Piston Cup Championship in California to compete against two seasoned pros, McQueen gets to know the town's offbeat characters.
Guido discovers he has a hidden talent as a street corner sign spinner.
A routine tow lands Mater in Tokyo, where he is challenged to a drift-style race against a nefarious gang leader and his posse of ninjas. With the help of his friend, 'Dragon' Lightning McQueen, and some special modifications, Mater attempts to drift to victory and become Tow-ke-O Mater, King of all Drifters.
Mater, the rusty but trusty tow truck from Cars, spends a day in Radiator Springs playing scary pranks on his fellow townsfolk. That night at Flo's V8 Café, the Sheriff tells the story of the legend of the Ghostlight, and as everyone races home Mater is left alone primed for a good old-fashioned scare.