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Chen Zhen, a Chinese engineering student in Kyoto, who braves the insults and abuse of his Japanse fellow students for his local love Mitsuko Yamada, daughter of the director, returns in 1937 to his native Shangai, under Japanse protectorate -in fact military occupation- after reading about the death of his kung-fu master Hou Ting-An in a fight against the Japanese champion Ryuichi Akutagawa.
With the Gyaos re-emerging, Gamera's ties to humanity have been severed with his bond to Asagi broken. Nagamine and Asagi investigate while an orphaned girl named Ayana discovers a new creature she names Iris. Nagamine and Asagi must reach Ayana before she takes her revenge on Gamera, who she blames for the death of her family.
The U.N.G.C.C. (United Nations Godzilla Countermeasure Center) recovers the remains of Mecha-King Ghidorah and construct Mechagodzilla as a countermeasure against Godzilla. Meanwhile, a giant egg is discovered along with a new monster called Rodan. The egg is soon found to be none other than an infant Godzillasaurus.
