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A submarine expedition to salvage the remains of Mechagodzilla is thwarted by a massive dinosaur named Titanosaurus. An Interpol investigation leads biologist Ichinose to uncover the work of Dr. Mafune and his mysterious daughter Katsura. Aligned with the Black Hole Aliens, Katsura's life becomes entwined with the resurrected machine.
Manga artist Gengo Odaka lands a job with the World Children's Land amusement park only to become suspicious of the organization when a garbled message is discovered on tapes. As Gengo and his team investigate, Godzilla and Anguirus quickly decipher the message and begin their own plan of action.
Hoping to find a sense of connection to her late mother, Gorgeous takes a trip to the countryside to visit her aunt at their ancestral house. She invites her six friends, Prof, Melody, Mac, Fantasy, Kung Fu, and Sweet, to join her. The girls soon discover that there is more to the old house than meets the eye.
An Okinawan prophecy that foretells the destruction of the Earth is seeming fulfilled when Godzilla emerges to return to his destructive roots. But not all is what it seems after Godzilla breaks his ally Anguirus's jaw. Matters are further complicated when a second Godzilla emerges, revealing the doppelgänger as a mechanical weapon.