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In an alternate timeline the original Godzilla is never defeated and repeatedly reemerges to feed on Japan's energy sources. A new inter-dimensional weapon called the Dimension Tide is created with the intent of eliminating Godzilla. However, the new weapon might also serve as a gateway to something far more sinister.
A strange meteor lands in Japan and unleashes hundreds of insect-like "Legion" creatures bent on colonizing the Earth. When the military fails to control the situation, Gamera shows up to deal with the ever-evolving space adversary. However the battle may result in Gamera losing his bond with both Asagi and humanity.
